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Encyclopedia Britannica



THORN (O. Eng. porn, cf. Du. doorn, Ger. Dorn, &c)

This article appears in Volume V26, Page 880 of the Encyclopedia Britannica.

Encyclopedia Britannica - Main :: THE-TOO
THORN (O. Eng. porn, cf. Du. doorn, Ger. Dorn, &c) , in botany, a hard pointed structure; also termed a " spine," generally representing a small branch, as in hawthorn, where a normal branch arising in the axil of a
leaf
  is replaced by a sharply pointed thorn; accessory buds on each side of the thorn and developed in the same
leaf
 -axil will grow in the next season into
ordinary
  branches. The similarly developed thorns of the honey-locust (Gleditschia) are branched. In other cases, as the sloe or the wild pear, branches become spiny at the apex tapering into a stiff leafless point. On a cultivated tree these branches disappear owing to their more vigorous growth. Leaves may be modified into spines, as in barberry, the leaves of which show every gradation between a leaf with a spiny-toothed edge and those which have been reduced to simple or multiple spines. In some species of Astrdgalus the petiole of the pinnately compound leaf persists after the fall of the leaflets as a
sharp
  spine. In the false acacia (
Robinia
 ) the stipules are represented by spines.
The reduction of the leaf-surface, of which the spinous habit is often an expression, is associated with growth in dry or exposed windy places. Thus, in the gorse, a characteristic plant of exposed localities such as open commons, the smaller branches, instead of being leaf-bearing shoots, are reduced to slender green spines, while the leaves on the main shoots are also more or less spinous in character. As the giving off of water from its surface is one of the
chief
  functions of a leaf, this process is thus reduced to a minimum in situations where water is scarce or would be liable to be given off too rapidly. An extreme case is afforded by the cacti and cactus-like euphorbias, which are a characteristic type of desert vegetation where water is extremely scarce. The whole plant is reduced to a simple or branching succulent, leafless, columnar or flattened
stem
 , the branches of which are represented by small clusters of thorns. Incidentally the
thorns protect the plant which bears them from the attacks of animals seeking food.
Prickles are structures of less importance from the morphological point of view, being mere superficial outgrowths which may occur anywhere on
stem
  or leaf, or even fruit.
THORNABY-ON-TEES, a municipal borough in the North Riding of Yorkshire, England, 3 m. S.W. of Middlesbrough, on the North-Eastern railway. Pop. (1901), 16,054. It lies opposite Stockton-on-Tees, with which it is connected by a bridge, on the river Tees. There are blast furnaces, iron foundries, engineering works, iron ship-building yards, extensive saw-mills, flour-mills and a manufactory of " blue and white " pottery. The town was formerly known as
South
  Stockton, and is still included in the parliamentary borough of Stockton (it is within the Cleveland division of the county), but was incorporated as a
separate
  municipal borough in 1892. It is under a mayor, 6 aldermen and 18 councillors. Area, 1927 acres.


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