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Encyclopedia Britannica



TABULAR VIEW OF

This article appears in Volume V10, Page 560 of the Encyclopedia Britannica.

Encyclopedia Britannica - Main :: SUS-TAV
TABULAR VIEW OF INFLORESCENCES
A. Indefinite Centripetal Inflorescence.
I.
Flowers
  solitary, axillary. Vinca, Veronica hederifolia.
II.
Flowers
  in groups, pedicellate.
1. Elongated form(Raceme), Hyacinth, Laburnum, Currant. (Corymb), Ornithogalum.
2. Contracted or shortened form (Umbel), Cowslip, Astrantia.
1. Elongated form (Spike), Plantago.
(Spikelet),
Grasses
 .
(Amentum, Catkin), Willow, Hazel.
(Spadix) Arum, some Palms.
(Strobilus), Hop.
2. Contracted or shortened fcrm (Capitulum), Daisy,Dandelion, Scabious.
IV. Compound Indefinite Inflorescence.
a. Compound Spike, Rye-grass.
b. Compound Spadix, Palms.
c. Compound Raceme, Astilbe,
d. Compound Umbel, Hemlock and most
Umbelliferae
 .
e. Raceme of Capitula, Petasites.
f. Raceme of Umbels, Ivy.
B. Definite Centrifugal Inflorescence.
I. Flowers solitary, terminal. Gentianella, Tulip.
II. Flowers in Cymes.
1. Uniparous Cyme.
a. Helicoid Cyme (axes forming a spiral).
Elongated form, Alstroemeria.
Contracted form, Witsenia corymbosa.
b. Scorpioid Cyme (axes unilateral, two rows). Elongated form, Forget-me-not, Symphylum, Henbane.
Contracted form, Erodium, Alchemilla arvensis. Biparous Cyme (Dichotomous),including 3-5-chotomous
Cymes (Dichasium, Cymose Umbel, Anthela).
a. Elongated form, Cerastium, Stellaria.
b. Contracted form (Verticillaster), Dead-nettle, Pelargonium.
3. Compound Definite Inflorescence. Streptocarpus polyanthus, many Calceolarias.
C. Mixed Inflorescence.
Raceme of Scorpioid Cymes, Horse-chestnut.
Scorpioid Cyme of Capitula, Vernonia scorpioides. Compound Umbel of Dichotomous Cymes,Laurustinus. Capitulum of contracted Scorpioid Cymes (Glomerulus),
Sea-
pink
 .
The flower consists of the floral
axis
  bearing the sporophylls
(stamens and carpels), usually with certain protective envelopes.
The
axis
  is usually very much
contracted, no inter-
nodes being devel-
oped, and the portion
bearing the floral leaves, termed
the thalamus or torus, frequently
expands into a conical, flattened
or hollowed expansion; at other
times, though rarely, the inter-
nodes are developed and it is
elongated. Upon this torus the
parts of the flower are arranged
in a crowded manner, usually
forming a
series
  of verticils, the
parts of which alternate; but
they are sometimes arranged
spirally especially if the floral
axis be elongated. In a typical
flower, as in fig. 22, we recognize
four distinct whorls of leaves:
an outer whorl, the calyx of
sepals; within it, another whorl,
the parts alternating with those
of the outer whorl, the corolla of
petals; next a whorl of parts
alternating with the parts of
the corolla, the androecium of
stamens; and in the centre the
gynoecium of carpels. Fig. 23 is
a diagrammatic representation
of the arrangement of the parts
of such a flower; it is known as
a floral diagram. The flower is
supposed to be cut transversely,
and the parts of each whorl
are distinguished by a different
symbol. Of these whorls the
two internal, forming the sporo-
phylls, constitute the essential
organs of reproduction; the two
outer whorls are the protective
coverings or floral envelopes. The
sepals are generally of a greenish
colour; their
function
  is mainly
protective, shielding the more
delicate internal organs before
the flower opens. The petals a-e
usually showy, and normally
alternate with the sepals. Some-
times, as usually in monocoty-
ledons, the calyx and corolla are
similar; in such cases the term
perianth, or perigone, is applied.
Thus, in the tulip, crocus, lily,
hyacinth, we speak of the parts of the perianth, in place of
calyx and corolla, although in these plants there is an outer
whorl (calyx), of three parts, and an inner (corolla), of a similar number, alternating with them. When the parts of the calyx are in
appearance
  like petals they are said to be petaloid, as in Liliaceae. In some cases the petals have the
appearance
  of sepals, then they are sepaloid, as in Juncaceae. In plants, as Nymphaea alba, where a spiral arrangement of the floral leaves occurs, it is not easy to say where the calyx ends and the corolla begins, as these two whorls pass insensibly into each other. When both calyx and corolla are present, the plants are dichlamydeous; when one only is present, the flower is termed monochlamydeous or apetalous, having no petals (fig. 24). Sometimes both are absent, when the flower is achlamydeous, or naked, as in willow. The outermost
series
  of the essential organs, collectively termed the androecium, is composed of the microsporophylls known as the staminal leaves or stamens. In their most differentiated form each consists of a stalk, the filament (fig. 25, f), supporting at its summit the anther (a), consisting of the pollen-sacs which contain the powdery pollen (p), the microspores, which is ultimately discharged therefrom. The gynoecium or pistil is the central portion of the flower, terminating the floral axis. It consists of one or more carpels (megasporophylls), either separate (fig. 22, c) or combined (fig. 24). The parts distinguished in the pistil, are the ovary (fig. 26, o), which is the lower portion enclosing the ovules destined to become seeds, and the stigma (g), a portion of loose cellular
tissue
 , the receptive surface on which the pollen is deposited, which is either sessile on the apex of the ovary, as in the poppy, or is separated from it by a prolonged portion called the style (s). The androecium and gynoecium are not present in all flowers. When both are present the flower is hermaphrodite; and in descriptive botany such a flower is indicated by the symbol . When only one of those organs is present the flower is unisexual or diclinous, and is either male (staminate),6, or female (pistillate), ~. A flower then normally consists of the four series of leavescalyx, corolla, androecium and gynoeciumand when these are all present the flower is complete. These are usually densely crowded upon the thalamus, but in some instances, after apical growth has ceased in the axis, an elongation of portions of 'the receptacle by intercalary growth occurs, by which changes in the position of the parts may be brought about. Thus in Lychnis an elongation of the axis betwixt the calyx and the corolla takes place, and in this way they are separated by an intervaL Again, in the passion-flower (Passiflora) the stamens are separated from the corolla by an elongated portion of the axis, which has consequently been termed the androphore, and in Passiflora also, fraxinella (fig. 27), Capparidaceae, and some other plants, the ovary is raised upon a distinct stalk termed the gynophore; it is thus separated from the stamens, and is said to be stipitate. Usually the successive whorls of the flower, disposed from below upwards or from without in-wards upon the floral axis, are of the same number of parts, or
are a multiple of the same number of parts, those of one whorl alternating with those of the whorls next it.
In the more primitive types of flowers the torus is more or less convex, and the series of organs follow in regular succession, culminating in the carpels, in the formation of which the growth of the axis is closed (fig. 28). This arrangement is known as hypogynous, the other series (calyx, corolla and stamens) beirg beneath (hypo-) the gynoecium. In other cases, the apex of the growing point ceases to develop, and the parts below form a cup around it, from the rim of which the outer members of the Power are developed around (peri-) the carpels, which are formed from the apex of the growing-point at the bottom of the cup. This arrangement is known as perigynous (fig. 29). In many cases this is carried farther and a cavity is formed which is roofed over
The flower.
and pistil of Fraxinella (Dictamnus Fraxinella). The pistil consists of several carpels, which are elevated on a stalk or gynophore prolonged from the receptacle.
by the carpels, so that the outer members of the flower
spring
  from the edge of the receptacle which is immediately above the ovary (epigynous), hence the term epigyny (fig. 30).


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