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Encyclopedia Britannica



ERICACEAE

This article appears in Volume V09, Page 740 of the Encyclopedia Britannica.

Encyclopedia Britannica - Main :: EMS-EUD
ERICACEAE , in botany, a natural order of plants belonging
to the higher or gamopetalous division of Dicotyledons. They
are woody plants, sometimes with a slender creeping stem as
in bilberry, Vaccinium (fig. 1), or Andromeda (fig. 2), or form-
ing low bushes as in
the heaths, or larger,
sometimes becoming
tree-like, as in species
of Rhododendron.
The leaves are alter-
nate, opposite or
whorled in arrange-
ment, and in their
form and structure
show well - marked
adaptation for life
in dry or exposed
situations. Thus in
the true. heaths they
are needle-like, with
the margins often
rolled back to form
a groove or an almost
closed chamber on
the under side. In
others such as Rhodo-
dendron or Arbutus
they are often
leathery and ever-
green, the strongly
cuticularized upper surface protecting a water-storing
tissue
 
situated above the green layers of the leaf. The
flowers
  are
sometimes solitary and axillary or terminal as in Andromeda,
but are generally arranged in racemose inflorescences at the end
of the branches as in Arbutus and Rhododendron, or on small
lateral shoots as in Erica. They are hermaphrodite and generally
regular with parts in 4 or 5, thus: sepals 4 or 5, petals 4 or. 5, stamens 8 or io in two
series
 , the outer of which is opposite the petals, and carpels 4 or 5. The corolla is usually more or less bell-shaped, and in the heaths persists in a dry state in the fruit. The petals with the stamens are situated on the outer edge of a
honey-secreting disk. The anthers show a very great variety in shape, the halves are often more or less free and often appendaged; they open to allow the escape of the pollen by a terminal pore or slit. The carpels are united to form a 4- to 5-chambered ovary, which bears a simple elongated style ending in a capitate stigma; each ovary-chamber contains one to many ovules attached to a central
placenta
 . The brightly coloured
1, Flowering shoot of Erica cinerea, about 12 nat. size.
2, Flower cut lengthwise. 5,
3, Stamen showing appendages
and porous dehiscence of
anther.
4, Capsule showing the loculicidal dehiscence ; a few seeds re-
corolla, the presence of
nectar
  and the scent render the
flowers
  attractive to insects, and the projection of the stigma beyond the anthers favours crossing. The fruit is generally a capsule containing many seeds, as in Erica (fig. 3) or Rhododendron; some-times a
berry
  as in Arbutus.
The order falls into four distinct tribes, which are characterized by the relative position of the ovary and by the fruit and seed. They are as follows:
1. Rhododendron tribe, characterized by capsular fruit, seed with a loose coat, deciduous petals and anthers without append-ages. It consists mainly of the great genus Rhododendron (in which Azalea is included by
recent
  botanists), which is chiefly
main attached to the central
axis
 .
Diagram of the flower having four sepals, four divisions of the corolla, eight stamens in two rows, and four divisions of the pistil.
developed in the mountains of eastern Asia, many species occur-ring on the Himalayas. Dabeocia, St Dabeoc's heath, occurs in Ireland.
2. Arbutus Tribe.Fruit a
berry
  or capsule, petals deciduous and anthers with bristle-like appendages, chiefly north temperate to arctic in distribution. Arbutus Unedo, the
strawberry
 -tree, so called from its large scarlet berry, is a southern European species which extends into south Ireland. Arctostaphylos (bearberry) and Andromeda are arctic and alpine genera occurring in Britain. Epigaea re pens is the trailing arbutus or mayflower of Atlantic America.
3. Vaccinium Tribe.Ovary inferior, fruit a berry. Extends from the north temperate zone to the mountains of the tropics. Vaccinium, the largest genus, has four British species: V. Myrtillus is the bilberry (q.v.), blaeberry or whortleberry, V. Vitis-Idaea the cowberry, and V. Oxycoccos the
cranberry
  (q.v.). This tribe is sometimes regarded as a separate order Vacciniaceae, distinguished by its inferior ovary.
4. Erica Tribe.Fruit usually a. capsule, seeds round, not winged; corolla persisting round the ripe fruit; anthers often appendaged. The largest genus is Erica, the true heath (q.v.), with over 400 species, the great majority of which are confined to the Cape; others occur on the mountains of tropical Africa and in Europe and North Africa, especially the Mediterranean region. E. cinerea (purple heather) and E. Tetralix (cross-leaved heath) are common British heaths. Calluna is the ling or Scotch heather.


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